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词条 瓣蕊花科
释义

§ 生物基本信息

瓣蕊花科[1] (HIMAMTANDRACEAE Diels 1917 nom. conserv.),大型、芳香的乔木,薄壁组织具球形油细胞,植物体含瓣蕊花碱(himandrin)和同类的生物碱(为一类特殊的吡啶类生物碱),但不含异喹啉类生物碱;一些细胞含草酸钙的单晶或簇晶;茎薄壁组织中分散有石细胞,在叶中也多少有分布;茎节具三叶隙;髓多少有良好发育的厚壁组织横隔;导管分子伸长,具单穿孔,或一部分在幼茎中具梯纹穿孔板;无穿孔的输水分子具具缘纹孔,不同的作者认为是管胞或纤维管胞;木薄壁组织为离管型,含晶体,并组成同心的带状;木射线细胞异形,或在晚材中近同形,大多数为(1)2-3(4)列;次生韧皮部在切向上分层,纤维层和非纤维层交替排列,并具楔状射线。叶互生,单叶,全缘,具透明腺点,羽状脉,下表面(及幼枝)覆盖有特征性的盾状毛;气孔为平列型,数个排成一环,每环各为一枚盾状毛遮掩;叶柄具环状排列的离生维管束;托叶缺。花相当大,单生(稀2-3)于侧枝上,两性,下位;萼片2,早落,帽状,一枚包住另一枚,外侧的一枚被认为是由2枚合生萼片演化而来,内侧的一枚则被认为是由4枚合生花瓣演化而来,或有时两者均被看作是苞片(如Endress);花冠由数枚(约7-9)螺旋状排列的披针状线形花瓣组成,这些花瓣毫无疑问是退化雄蕊起源的,如果把内侧的帽状萼片看成变态的花冠的话,那么这些花瓣就可解释为是退化雄蕊;雄蕊多数(约25-30),螺旋状排列,外观极似花瓣,不分化为花丝和花药。4个小孢子囊相当短,多少嵌生于片状雄蕊中部以下的远轴一边,在中脉和边脉间两两成对,每一对沿单独一缝开裂;数枚(约8-10)螺旋状排列的退化雄蕊生于正常雄蕊和雌蕊间;花粉粒近球形,具单沟(即远极沟),具多少坚实、均质的外壁;雌蕊约(6)7-10或多数(多至约28),螺旋状排列,心皮闭合,明显分化为子房和具下延柱头的短花柱,花期各心皮在基部微合生,果期则较充分地以边缘合生;每室具单1(稀为2)胚珠,悬垂,片状边缘胎座,倒生。果实球形,虫瘿状,肉质,因心皮的愈合而为多室;种子具小形、双子叶的胚和丰富、富含油脂、非嚼烂状的胚乳。2n=24。

§ 种族信息

瓣蕊花科仅含瓣蕊花属Galbulimina (Himantandra)一属,分布限于新几内亚、马鲁古(Molucca)群岛和澳大利亚东北部。一种观点认为这三个地区各有一特有种,另一种观点则认为只有一种,即瓣蕊花G. belgraveana (F. Muell.) Sprague。

瓣蕊花科一般认为和木兰科、单心木兰科和番荔枝科有联系。因此,即使在定义较狭的木兰目中,它们也仍被保留。不过,本科每心皮通常只具单1胚珠,这一特征在木兰目中比较特殊,而与樟目(Laurales)相似。和木兰目中其他各科一样,本科也代表了一个孤立进化死支,和其他各科均无密切联系。

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